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Geographic distribution of Ischyropsalis impressa sp. nov. (yellow dots) and I. aguerana sp. nov. (white dots) in northeastern Cantabrian Mountains. A) Location of study area and map of the Karst in <t>Spain</t> at 1:1,000,000-scale where karstified carbonate rocks emerge. B) LiDAR visualisation of the study area <t>showing</t> <t>Cantabria–Biscay</t> border (solid yellow line). The orange-brown area represents the range of limestone outcrops from the Urgonian formation (of Aptian age, Early Cretaceous) at 1:50,000-scale. Location of the caves surveyed: 1. La Baja; 2. Hoyo de Peñaflor; 3. Perelada; 4. Hoyo Molino; 5. Hoyo Menor-Tocinos; 6. Hoyo Menor-Rejuyo; 7. Hoyo de la Cubilla; 8. La Lastrilla; 9. Covacho del Cogorio; 10. Hoyo de las Rebolligas; 11. La Pasada; 12. Hoyo de las Fuentes; 13. La Tabla II; 14. Gueriza.
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Geographic distribution of Ischyropsalis impressa sp. nov. (yellow dots) and I. aguerana sp. nov. (white dots) in northeastern Cantabrian Mountains. A) Location of study area and map of the Karst in <t>Spain</t> at 1:1,000,000-scale where karstified carbonate rocks emerge. B) LiDAR visualisation of the study area <t>showing</t> <t>Cantabria–Biscay</t> border (solid yellow line). The orange-brown area represents the range of limestone outcrops from the Urgonian formation (of Aptian age, Early Cretaceous) at 1:50,000-scale. Location of the caves surveyed: 1. La Baja; 2. Hoyo de Peñaflor; 3. Perelada; 4. Hoyo Molino; 5. Hoyo Menor-Tocinos; 6. Hoyo Menor-Rejuyo; 7. Hoyo de la Cubilla; 8. La Lastrilla; 9. Covacho del Cogorio; 10. Hoyo de las Rebolligas; 11. La Pasada; 12. Hoyo de las Fuentes; 13. La Tabla II; 14. Gueriza.
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Geographic distribution of Ischyropsalis impressa sp. nov. (yellow dots) and I. aguerana sp. nov. (white dots) in northeastern Cantabrian Mountains. A) Location of study area and map of the Karst in <t>Spain</t> at 1:1,000,000-scale where karstified carbonate rocks emerge. B) LiDAR visualisation of the study area <t>showing</t> <t>Cantabria–Biscay</t> border (solid yellow line). The orange-brown area represents the range of limestone outcrops from the Urgonian formation (of Aptian age, Early Cretaceous) at 1:50,000-scale. Location of the caves surveyed: 1. La Baja; 2. Hoyo de Peñaflor; 3. Perelada; 4. Hoyo Molino; 5. Hoyo Menor-Tocinos; 6. Hoyo Menor-Rejuyo; 7. Hoyo de la Cubilla; 8. La Lastrilla; 9. Covacho del Cogorio; 10. Hoyo de las Rebolligas; 11. La Pasada; 12. Hoyo de las Fuentes; 13. La Tabla II; 14. Gueriza.
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Geographic distribution of Ischyropsalis impressa sp. nov. (yellow dots) and I. aguerana sp. nov. (white dots) in northeastern Cantabrian Mountains. A) Location of study area and map of the Karst in <t>Spain</t> at 1:1,000,000-scale where karstified carbonate rocks emerge. B) LiDAR visualisation of the study area <t>showing</t> <t>Cantabria–Biscay</t> border (solid yellow line). The orange-brown area represents the range of limestone outcrops from the Urgonian formation (of Aptian age, Early Cretaceous) at 1:50,000-scale. Location of the caves surveyed: 1. La Baja; 2. Hoyo de Peñaflor; 3. Perelada; 4. Hoyo Molino; 5. Hoyo Menor-Tocinos; 6. Hoyo Menor-Rejuyo; 7. Hoyo de la Cubilla; 8. La Lastrilla; 9. Covacho del Cogorio; 10. Hoyo de las Rebolligas; 11. La Pasada; 12. Hoyo de las Fuentes; 13. La Tabla II; 14. Gueriza.
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Geographic distribution of Ischyropsalis impressa sp. nov. (yellow dots) and I. aguerana sp. nov. (white dots) in northeastern Cantabrian Mountains. A) Location of study area and map of the Karst in <t>Spain</t> at 1:1,000,000-scale where karstified carbonate rocks emerge. B) LiDAR visualisation of the study area <t>showing</t> <t>Cantabria–Biscay</t> border (solid yellow line). The orange-brown area represents the range of limestone outcrops from the Urgonian formation (of Aptian age, Early Cretaceous) at 1:50,000-scale. Location of the caves surveyed: 1. La Baja; 2. Hoyo de Peñaflor; 3. Perelada; 4. Hoyo Molino; 5. Hoyo Menor-Tocinos; 6. Hoyo Menor-Rejuyo; 7. Hoyo de la Cubilla; 8. La Lastrilla; 9. Covacho del Cogorio; 10. Hoyo de las Rebolligas; 11. La Pasada; 12. Hoyo de las Fuentes; 13. La Tabla II; 14. Gueriza.
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holotype  (Brinkmann Instruments)
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Geographic distribution of Ischyropsalis impressa sp. nov. (yellow dots) and I. aguerana sp. nov. (white dots) in northeastern Cantabrian Mountains. A) Location of study area and map of the Karst in <t>Spain</t> at 1:1,000,000-scale where karstified carbonate rocks emerge. B) LiDAR visualisation of the study area <t>showing</t> <t>Cantabria–Biscay</t> border (solid yellow line). The orange-brown area represents the range of limestone outcrops from the Urgonian formation (of Aptian age, Early Cretaceous) at 1:50,000-scale. Location of the caves surveyed: 1. La Baja; 2. Hoyo de Peñaflor; 3. Perelada; 4. Hoyo Molino; 5. Hoyo Menor-Tocinos; 6. Hoyo Menor-Rejuyo; 7. Hoyo de la Cubilla; 8. La Lastrilla; 9. Covacho del Cogorio; 10. Hoyo de las Rebolligas; 11. La Pasada; 12. Hoyo de las Fuentes; 13. La Tabla II; 14. Gueriza.
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Image Search Results


Geographic distribution of Ischyropsalis impressa sp. nov. (yellow dots) and I. aguerana sp. nov. (white dots) in northeastern Cantabrian Mountains. A) Location of study area and map of the Karst in Spain at 1:1,000,000-scale where karstified carbonate rocks emerge. B) LiDAR visualisation of the study area showing Cantabria–Biscay border (solid yellow line). The orange-brown area represents the range of limestone outcrops from the Urgonian formation (of Aptian age, Early Cretaceous) at 1:50,000-scale. Location of the caves surveyed: 1. La Baja; 2. Hoyo de Peñaflor; 3. Perelada; 4. Hoyo Molino; 5. Hoyo Menor-Tocinos; 6. Hoyo Menor-Rejuyo; 7. Hoyo de la Cubilla; 8. La Lastrilla; 9. Covacho del Cogorio; 10. Hoyo de las Rebolligas; 11. La Pasada; 12. Hoyo de las Fuentes; 13. La Tabla II; 14. Gueriza.

Journal: bioRxiv

Article Title: Unraveling the Phylogenetic Complexities of Ischyropsalis C.L. Koch, 1839 (Opiliones: Dyspnoi: Ischyropsalididae) in the North Iberian Peninsula, with the description of three new species

doi: 10.64898/2026.01.09.698449

Figure Lengend Snippet: Geographic distribution of Ischyropsalis impressa sp. nov. (yellow dots) and I. aguerana sp. nov. (white dots) in northeastern Cantabrian Mountains. A) Location of study area and map of the Karst in Spain at 1:1,000,000-scale where karstified carbonate rocks emerge. B) LiDAR visualisation of the study area showing Cantabria–Biscay border (solid yellow line). The orange-brown area represents the range of limestone outcrops from the Urgonian formation (of Aptian age, Early Cretaceous) at 1:50,000-scale. Location of the caves surveyed: 1. La Baja; 2. Hoyo de Peñaflor; 3. Perelada; 4. Hoyo Molino; 5. Hoyo Menor-Tocinos; 6. Hoyo Menor-Rejuyo; 7. Hoyo de la Cubilla; 8. La Lastrilla; 9. Covacho del Cogorio; 10. Hoyo de las Rebolligas; 11. La Pasada; 12. Hoyo de las Fuentes; 13. La Tabla II; 14. Gueriza.

Article Snippet: Type material Holotype SPAIN – Cantabria Province • ♂; Guriezo; La Pasada Cave; 30TVN7689794385, 440 m a.s.l.

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